Fittings are just a restriction and flow is NOT "limited" by them... in the sense, 1/4" air line with 1/4" fittings = 3/8" line with 1/4" fitting, is false in common air setups.
Each fitting add to the overall restriction. The air line is usually the greatest restriction, due to length. 3/8" air line with 1/4" fittings is generally sufficient for most automotive tasks: impact, ratchet, die grinder, etc... Whereas for a carpenter, a 1/4" air line would be fine with nail guns. Someone in heavy industry running 3/4"+ impacts and blast cabinets, might need 1/2" air lines.